region      reef                                           lat      lon      database_id  locus  ancestral_allele_1  ancestral_allele_2  allele_3  allele_4  origin_allele_3  origin_allele_4  mutation_step_allele_3  mutation_step_allele_4  mutational_change_allele_3_bp  mutational_change_allele_4_bp  sample_count_for_genet  genet_id  full_mutation  
Panama      Bastimentos                                    nd       nd       2003         192    163                 166                 160       nd        1                nd               1                       nd                      -3                             nd                             16                      P1150     nd             
Panama      Bastimentos                                    nd       nd       2047         182    155                 161                 158       nd        2                nd               1                       nd                      -3                             nd                             3                       P1166     nd             
Panama      Bocas_Del_Drago                                nd       nd       2061         192    154                 193                 190       196       2                2                1                       1                       -3                             3                              5                       P1167     nd             
Panama      Bocas_Del_Drago                                nd       nd       2068         192    154                 193                 196       nd        2                nd               1                       nd                      3                              nd                             5                       P1167     nd             
Panama      Bocas_Del_Drago                                nd       nd       2049         207    173                 191                 188       nd        2                nd               1                       nd                      -3                             nd                             5                       P1167     nd             
Panama      Bocas_Del_Drago                                nd       nd       2066         182    173                 176                 179       nd        2                nd               1                       nd                      3                              nd                             15                      P1168     nd             
Panama      Bocas_Del_Drago                                nd       nd       2057         192    166                 166                 169       nd        CBD              nd               nd                      nd                      nd                             nd                             15                      P1168     nd             
Panama      Wild_Cayne                                     nd       nd       2079         192    172                 175                 178       nd        2                nd               1                       nd                      3                              nd                             10                      P1177     nd