region      reef                                           lat      lon      database_id  locus  ancestral_allele_1  ancestral_allele_2  allele_3  allele_4  origin_allele_3  origin_allele_4  mutation_step_allele_3  mutation_step_allele_4  mutational_change_allele_3_bp  mutational_change_allele_4_bp  sample_count_for_genet  genet_id  full_mutation  
Panama      Bastimentos                                    nd       nd       2003         192    163                 166                 160       nd        1                nd               1                       nd                      -3                             nd                             16                      P1150     nd             
Panama      Bastimentos                                    nd       nd       2047         182    155                 161                 158       nd        2                nd               1                       nd                      -3                             nd                             3                       P1166     nd